_a b n_ is the membrane which closes the eye from below, upwards,
with an opaque film, _c n b_ encloses the eye in front and behind
with a transparent membrane.

It closes from below, upwards, because it [the eye] comes downwards.

When the eye of a bird closes with its two lids, the first to close
is the nictitating membrane which closes from the lacrymal duct over
to the outer corner of the eye; and the outer lid closes from below
upwards, and these two intersecting motions begin first from the
lacrymatory duct, because we have already seen that in front and
below birds are protected and use only the upper portion of the eye
from fear of birds of prey which come down from above and behind;
and they uncover first the membrane from the outer corner, because
if the enemy comes from behind, they have the power of escaping to
the front; and again the muscle called the nictitating membrane is
transparent, because, if the eye had not such a screen, they could
not keep it open against the wind which strikes against the eye in
the rush of their rapid flight. And the pupil of the eye dilates and
contracts as it sees a less or greater light, that is to say intense
brilliancy.